NATURE

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Title:  Rapture of the Deep	
Authors:  Michael Tennesen
Source:  WILDLIFE CONSERVATION; May/June 2005	
Database:  D Drive


RAPTURE OF THE DEEP

By Michael Tennesen

During the austral summer, the human population at McMurdo Station on Ross 
Island just off the Antarctic shore swells from around 250 to more than 1,000 
as the scientists take advantage of 24-hour sunlight and "toasty," 
near-freezing temperatures. When Gerald Kooyman, a research professor at 
Scripps Institution of Oceanography in San Diego, California, first visited 
the station in the 1960s, he looked out across McMurdo Sound, a sheet of thick 
ice wedged between Ross Island and the continent, and wondered what he could 
do with this great outdoor laboratory. With no natural predators, the animals 
were unafraid of people, so he decided to elicit their help in a diving 
experiment. Kooyman transplanted a Weddell seal out to a hole he had drilled 
with a large augur into the six-foot thick ice. The hole was placed far away 
from other holes or open ice so that the seal would use it to both dive and 
to return to the surface. Thus the scientist wouldn't lose the expensive 
diving recorder he'd attached to the seal, and no one could claim the seals 
went somewhere else during the experiment.

A pioneer in the study of deep diving, Kooyman began his work at a time when 
scientists generally thought that animals might be able to descend 325 to 650 
feet below the surface. "I remember I was eating a dry cracker when the depth 
recorder came back from one seal that showed it dove to 1,970 feet," he 
recalls. "I nearly choked on the cracker."

A lot of scientists have since "gagged on their crackers," and called 
manufacturers to double-check the accuracy of their depth recorders as, one 
after another, the emperor penguin, leatherback sea turtle, northern elephant 
seal, bottlenose whale, and sperm whale has met and surpassed that record. 
Emperor penguins can dive to 1,853 feet, and sperm whales to 6,560 feet

Compare those depths to that of humans. The world record for a human descent 
without diving gear but assisted by ropes and pulleys is 531 feet. And human 
divers attempting these depths and greater, with or without gear, are subject 
to a host of maladies including shallow-water blackouts (the sudden loss of 
consciousness caused by oxygen starvation following a breath-holding dive) 
and the bends, or decompression sickness-both of which can be fatal. But 
deep-diving animals glide on through water levels dangerous to people without 
even elevated heart rates.

Kooyman learned more about the ease with which deep-diving animals accomplish 
these phenomenal dives when he returned to McMurdo more recently to test 
emperor penguins. For a number of years, Kooyman and his associate Paul 
Ponganis, a research physiologist at Scripps and a practicing 
anesthesiologist, have been trying to understand some of the remarkable 
physiological adaptations that enable these animals to accomplish their 
remarkable diving feats.

For better than two weeks, they worked with ten adult penguins, each of which 
weighed from 45 to 60 pounds. The scientists divided the birds into two 
groups: one group they fed by hand, and the other was allowed to hunt its own 
prey through a hole drilled in the ice. By taking blood samples from the two 
groups and comparing the results, the scientists hoped to measure the 
energetic costs of diving for prey.

But when the results come back they were shocked. Emperor penguins chasing 
prey in frigid Antarctic waters are such efficient predators that they don't 
use any more energy than if they were simply standing around, chowing down on 
handouts. "For penguins and seals, hunting is more like meditation," says 
Ponganis,

To get into that state, Weddell seals and emperor penguins exhibit an 
exquisite ability to control their own heart rates during dives. Normal 
heart rates for emperor penguins are 60 to 70 beats per minute; but when the 
birds dive, the heart rate can go as low as 20 to 30. They essentially save 
most of their bodily functions-purging carbon dioxide, eliminating waste, 
and even digesting food-for the surface, when their heart rates may go up 180 
to 200 beats per minute. During deep dives, the emperor's body may even shut 
down organs such as the liver, kidneys, and stomach.

In elephant seals, the heart rate of at the surface is 120 beats per minute; 
at depths, it falls to 30 to 35 beats, and has been recorded as low as 2 
beats per minute. That's 30 seconds between pulses-approaching cardiac 
arrest in a human. Unlike in humans, most of the oxygen to sustain the 
seals' long, deep dives is stored in blood and muscle, not in the lungs. 
Oxygen is bound to the protein myoglobin in the muscles and hemoglobin in 
the blood. Deep divers have a high concentration of hemoglobin in their 
blood and have a large blood volume. With all this oxygen already aboard, 
they don't need to work themselves into a heavy breathing frenzy, like land 
animals do, when they are chasing prey. 

What emperor penguins, Weddell seals, and elephant seals share with a number 
of other deep divers are streamlined bodies and enough mass that they glide 
through the water with little effort. During a week on the Island of San 
Miguel, off the coast of California, we witnessed how incredibly awkward 
elephant seals are on land-lurching across the sand as one scientist put it 
like "enormous fat caterpillars."

But when one male elephant seal hauled himself from the shore into the water, 
he metamorphosed into a ballet-like swimmer, instantly attaining speed while 
displaying little effort. Says Michael Castelini, professor of marine science 
at the University of Alaska at Fairbanks, "We [humans] thrash and flail our 
way through the water, while the elephant seal glides through it as if it 
were on a layer of ball bearings."

Diving may be effortless for elephant seals. Biologists have recorded one 
seal diving to 5,479 feet-a record for the species. To put that in 
perspective, imagine more than three Empire State buildings stacked on top of 
one another. The seal swam from the top of the spire of the uppermost 
building to the basement of the bottom building and back up again, a distance 
of over two miles, in less than an hour.

There are animals that live at such depths, such as the coelecanth, but they 
often die when pulled to the surface, or live for a only very short time out 
of their element. Elephant seals can survive at depth and at the surface, 
though these environments are as different as those on Mars and Venus.

What's more, when elephant seals come to the surface, they spend only a few 
minutes there, exhaling carbon dioxide and replenishing oxygen, before they 
dive again. According to Robert DeLong, research biologist for the National 
Marine Mammal Laboratory in Seattle, "You want to think of them not as a 
diving animal but as an animal that occasionally surfaces. They spend 90 
percent of their time at sea under water."

And they can stay under for long periods without breathing. One female 
tagged by DeLong, dove for two hours.

For part of that time, the animal may be sleeping. Diving may be almost an 
auto pilot effort, as the seals take short naps on the way down and up. 
Castellini believes that elephant seals may sleep as dolphins do: closing 
one eye while half of the brain dozes and the other half keeps vigilance, 
then switching back and forth. If elephant seals can go deep enough, they 
don't have to worry about predation by great white sharks or killer 
whales, so swimming at reduced vigilance may not be a problem.

But how do elephant seals avoid rapture of the deep-a common name for 
decompression sickness? Researchers believe that an elephant seal's lungs 
collapse at a depths between 165 and 230 feet. With no exchange of gases 
(particularly nitrogen) between lungs and blood, deep-diving animals avoid 
the blood-chemistry imbalances that create human diving problems.

Still they must survive enormous pressures. Every 10 meters a deep diver 
descends, the pressure increases by approximately one atmosphere. That means 
for a sperm whale to dive 6,560 feet, it must endure pressures equal to about 
194 atmospheres or 2845 pounds on every square inch of its skin. 

Says DeLong, "Can you imagine what they must look like? Clearly the whole 
chest cavity collapses and the animal's face must look like a prune."

Students of Le Boeuf like to paint up Styrofoam mannequins, put lipstick on 
them, color their eyes, and then send them down 300 feet, just for fun. 
According to Le Boeuf, "They come back looking like shrunken heads."

The first recording devices that were placed on elephant seals were metal 
boxes, and they imploded from the pressure.

The speeds at which deep-diving animals make their descents are another 
marvel. Says Ponganis, "I talk to physicians who manage experimental human 
divers, and they are absolutely floored at how fast seals and whales are 
diving. Safely taking a man down to 1,900 feet can take four days, whereas 
an emperor penguin can get there in five minutes."

But why go there? What is the advantage? Many of the deep divers are going 
to an area called the deep scattering layer, a horizontal zone of living 
organisms, usually schools of fish, occurring below the surface in many ocean 
areas. The name comes from the fact that this layer scatters or reflects 
sound waves. This rich zone contains hundreds of species of fish, which 
migrate closer to the surface at night but retreat to the depths by day. 
Says Le Boeuf, "The deep scattering layer is where most of the biomass in 
the ocean is concentrated. These seals are diving to the center of the 
richest part."

By venturing to a zone where few large competitors go, the seals have a 
virtual monopoly on some of the richest waters off the continental shelf. 
And spending very little time at the surface, they avoid two deadly 
predators, the great white shark and the killer whale, which patrol 
shallower waters.

The deepest diver may not be a seal or a penguin, but a whale. The sperm 
whale, the reigning dive champion, has been recorded at depths of 6,650 
FEET. Even leatherback sea turtles have been recorded at depths below 
3,280 FEET. However, Sascha Hooker, a biologist with the University of St. 
Andrews in the United Kingdom, thinks that if you are interested in what 
animals actually do for a living, then looking at their deepest dives is 
not as important as looking at their routine dives. Sperm whales routinely 
dive to 1,300 to 1,950 feet , elephant seals normally dive to 1,150 to 
1,950 feet, and bottlenose whales, which Hooker studied in a submarine 
canyon off Nova Scotia, regularly make dives to 3,300 to 4,900 feet. At 
these levels there is little or no light. Cameras attached to animals that 
go to these depths return with images of a black screen 90 percent of the 
time.

Hal Whitehead, a biologist at Dalhousie University in Halifax, Nova Scotia, 
says that some fish at these depths are bioluminescent, and whales and 
seals may key on this bodily light. Or they may swim beneath their prey in 
daylight hours and look upward for silhouettes; hence, the elephant seals' 
enormous eyes help them see in the darkness at depths. Many of the whales 
use natural sonar systems to locate prey. The nose of the sperm whale, 
which Whitehead studies, constitutes a quarter to a third of its total 
weight. He thinks that nose contains the most powerful sonar system in 
the natural world.

He also thinks that the greatest advantage of diving to depth for a living is 
the speed, or lack thereof, of the prey down there. "Animals at the surface 
tend to be fast or fierce. This is possible because surface waters have a lot 
of oxygen. But when you get down below 1,300 feet, there is very little 
oxygen, and the escape responses of prey are very limited."

What that means is if you can handle the crushing pressure, don't get the 
bends, hold your breath for ten minutes to an hour, have huge eyes or a sonar 
system built into your nose, then the deep sea could just be one of the 
biggest all-you-can eat buffets the Earth has to offer.

                               -- End --

Freelance writer Michael Tennesen last wrote about thieving crows in our 
April 2005 issue.